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(2001). Cognition as expression: on the autopoietic foundations of an aesthetic theory of nature. Sign systems studies 29 (1), pp. 153-167.

(2002). Feeling the signs: the origins of meaning in the biological philosophy of Susanne K. Langer and Hans Jonas. Sign systems studies 30 (1), pp. 183-199.

(2002). Tundes märke: tähenduse päritolu susanne k. Langeri ja Hans Jonase bioloogilises filosoofias. kokkuvõte. Sign systems studies 30 (1), pp. 199-200.

with Varela, F. (2002). Life after Kant: natural purposes and the autopoietic foundations of biological individuality. Phenomenology and the Cognitive Sciences 1 (2), pp. 97-125.

(2004). Mimesis and metaphor: the biosemiotic generation of meaning in Cassirer and Uexküll. Sign systems studies 32 (1-2), pp. 297-307.

(2015)., Die Wiedergefundene Welt, in B. Pörksen (Hrsg.), Schlüsselwerke des Konstruktivismus, Wiesbaden, Verlag für Sozialwissenschaften, pp. 291-308.